:root {
  --color-ivory: #f5f1eb;
  --color-paper: #fffdf9;
  --color-ink: #25221f;
  --color-muted: #6e6861;
  --color-terracotta: #8f3f30;
  --color-sage: #2f634e;
  --color-line: #d8d0c7;
  --color-soft-gray: #e6e0d8;
  --font-display: Georgia, 'Times New Roman', serif;
  --font-body: Inter, ui-sans-serif, system-ui, sans-serif;
  --shadow-soft: 0 18px 50px rgba(37, 34, 31, 0.08);
  --radius-card: 4px;
}

*,
*::before,
*::after {
  box-sizing: border-box;
}

html {
  scroll-behavior: smooth;
  background: var(--color-ivory);
}

body {
  margin: 0;
  color: var(--color-ink);
  background: var(--color-ivory);
  font-family: var(--font-body);
  font-size: 1rem;
  line-height: 1.6;
  overflow-x: hidden;
}

body,
button,
a {
  -webkit-font-smoothing: antialiased;
}

a {
  color: inherit;
}

button,
a {
  -webkit-tap-highlight-color: transparent;
}

.container {
  width: min(1120px, calc(100% - 5rem));
  margin-inline: auto;
}

.sr-only {
  position: absolute;
  width: 1px;
  height: 1px;
  padding: 0;
  margin: -1px;
  overflow: hidden;
  clip: rect(0, 0, 0, 0);
  white-space: nowrap;
  border: 0;
}

.skip-link {
  position: fixed;
  z-index: 20;
  top: 0.75rem;
  left: 1rem;
  padding: 0.7rem 1rem;
  color: var(--color-paper);
  background: var(--color-ink);
  transform: translateY(-180%);
  transition: transform 160ms ease;
}

.skip-link:focus {
  transform: translateY(0);
}

.site-header {
  position: sticky;
  z-index: 10;
  top: 0;
  border-bottom: 1px solid var(--color-line);
  background: rgba(245, 241, 235, 0.94);
  backdrop-filter: blur(14px);
}

.site-header__inner {
  display: flex;
  min-height: 5.7rem;
  align-items: center;
  justify-content: space-between;
  gap: 2rem;
}

.site-header__identity,
.site-footer__identity {
  display: flex;
  align-items: center;
  gap: 1.25rem;
}

.brand {
  display: inline-flex;
  align-items: center;
  gap: 0.8rem;
  color: var(--color-ink);
  font-weight: 700;
  letter-spacing: 0.12em;
  text-decoration: none;
}

.brand__mark {
  display: grid;
  width: 2.45rem;
  height: 2.45rem;
  place-items: center;
  border: 1px solid var(--color-terracotta);
  border-radius: 50%;
  color: var(--color-terracotta);
  font-size: 1.45rem;
  line-height: 1;
}

.brand__name {
  font-size: 1.2rem;
}

.institutional-mark {
  display: inline-flex;
  align-items: center;
  gap: 0.55rem;
  color: var(--color-muted);
  font-size: 0.68rem;
  font-weight: 700;
  letter-spacing: 0.08em;
  line-height: 1.2;
  text-decoration: none;
  text-transform: uppercase;
}

.institutional-mark img {
  display: block;
  flex: 0 0 auto;
  width: 4.4rem;
  height: 1.35rem;
  object-fit: cover;
  object-position: center;
  mix-blend-mode: multiply;
}

.institutional-mark:hover,
.institutional-mark:focus-visible {
  color: var(--color-terracotta);
}

.institutional-mark--header {
  padding-left: 1.25rem;
  border-left: 1px solid var(--color-line);
}

.institutional-mark--section {
  margin-bottom: 1.4rem;
}

.section-heading__row {
  display: flex;
  align-items: center;
  justify-content: space-between;
  gap: 2rem;
}

.section-heading__row .eyebrow {
  margin-bottom: 0;
}

.site-nav {
  display: flex;
  flex-wrap: wrap;
  align-items: center;
  justify-content: flex-end;
  gap: 0.35rem 1.2rem;
}

.site-nav a,
.text-link {
  color: var(--color-muted);
  font-size: 0.86rem;
  font-weight: 700;
  letter-spacing: 0.02em;
  text-decoration: none;
}

.site-nav a {
  padding: 0.5rem 0;
}

.site-nav a:hover,
.site-nav a:focus-visible,
.text-link:hover,
.text-link:focus-visible {
  color: var(--color-terracotta);
}

.menu-toggle {
  display: none;
  min-width: 2.8rem;
  min-height: 2.8rem;
  border: 1px solid var(--color-line);
  border-radius: 50%;
  color: var(--color-ink);
  background: var(--color-paper);
  font-size: 1.2rem;
  cursor: pointer;
}

.section {
  position: relative;
  isolation: isolate;
  padding: clamp(6rem, 11vw, 10rem) 0;
  scroll-margin-top: 5.7rem;
  overflow: hidden;
}

.section--hero {
  min-height: min(860px, calc(100svh - 5.7rem));
  display: flex;
  flex-direction: column;
  justify-content: center;
  padding-top: clamp(4.5rem, 8vw, 7rem);
  padding-bottom: 3rem;
}

.hero__grid {
  display: grid;
  grid-template-columns: minmax(0, 1.05fr) minmax(300px, 0.95fr);
  align-items: center;
  gap: clamp(3rem, 8vw, 8rem);
}

.hero__copy {
  position: relative;
  z-index: 1;
  max-width: 44rem;
}

.eyebrow {
  margin: 0 0 1.35rem;
  color: var(--color-terracotta);
  font-size: 0.82rem;
  font-weight: 700;
  letter-spacing: 0.16em;
  line-height: 1.35;
  text-transform: uppercase;
}

h1,
h2,
h3,
p {
  margin-top: 0;
}

h1,
h2,
h3 {
  font-family: var(--font-display);
  font-weight: 400;
  line-height: 1.02;
}

h1 {
  max-width: 12ch;
  margin-bottom: 2rem;
  font-size: clamp(3.4rem, 6.8vw, 6.3rem);
  letter-spacing: -0.055em;
}

h2 {
  max-width: 14ch;
  margin-bottom: 1.5rem;
  font-size: clamp(2.6rem, 5vw, 5rem);
  letter-spacing: -0.045em;
}

h3 {
  margin-bottom: 0.7rem;
  font-size: clamp(1.4rem, 2vw, 1.9rem);
  letter-spacing: -0.025em;
}

.hero__lede {
  max-width: 39rem;
  margin-bottom: 2.25rem;
  color: var(--color-muted);
  font-size: clamp(1.05rem, 1.7vw, 1.3rem);
}

.button-row {
  display: flex;
  flex-wrap: wrap;
  align-items: center;
  gap: 0.8rem;
}

.button {
  display: inline-flex;
  min-height: 3.55rem;
  align-items: center;
  justify-content: center;
  gap: 0.7rem;
  padding: 0.85rem 1.45rem;
  border: 1px solid transparent;
  border-radius: 999px;
  font-weight: 700;
  text-decoration: none;
  transition: transform 180ms ease, background-color 180ms ease, border-color 180ms ease;
}

.button:hover {
  transform: translateY(-2px);
}

.button--primary {
  color: var(--color-paper);
  background: var(--color-terracotta);
}

.button--primary:hover,
.button--primary:focus-visible {
  background: #773326;
}

.button--quiet {
  border-color: var(--color-line);
  color: var(--color-ink);
  background: transparent;
}

.button--quiet:hover,
.button--quiet:focus-visible {
  border-color: var(--color-terracotta);
  background: rgba(255, 253, 249, 0.7);
}

.hero__signal {
  position: relative;
  display: grid;
  width: min(100%, 29rem);
  aspect-ratio: 1;
  margin-inline: auto;
  place-items: center;
  color: var(--color-ink);
}

.signal-orbit {
  position: absolute;
  border: 1px solid var(--color-line);
  border-radius: 50%;
  animation-timing-function: linear;
  animation-iteration-count: infinite;
}

.signal-orbit::before,
.signal-orbit::after {
  position: absolute;
  width: 0.8rem;
  height: 0.8rem;
  border-radius: 50%;
  content: '';
}

.signal-orbit::before {
  top: 2%;
  right: 8%;
  background: var(--color-terracotta);
}

.signal-orbit::after {
  bottom: 14%;
  left: 3%;
  background: var(--color-sage);
}

.signal-orbit--outer {
  width: 100%;
  height: 100%;
  animation: signal-spin-outer 28s linear infinite;
}

.signal-orbit--middle {
  width: 76%;
  height: 76%;
  border-color: var(--color-sage);
  animation: signal-spin-middle 23s linear infinite reverse;
}

.signal-orbit--inner {
  width: 52%;
  height: 52%;
  border-color: rgba(143, 63, 48, 0.35);
  animation: signal-spin-inner 17s linear infinite;
}

.signal-core {
  position: relative;
  z-index: 1;
  display: grid;
  width: 47%;
  height: 47%;
  place-content: center;
  gap: 0.15rem;
  border-radius: 50%;
  color: var(--color-paper);
  background: var(--color-terracotta);
  box-shadow: 0 22px 48px rgba(143, 63, 48, 0.23);
  font-size: clamp(0.63rem, 1.2vw, 0.85rem);
  font-weight: 700;
  letter-spacing: 0.18em;
  text-align: center;
}

.signal-core strong {
  color: rgba(255, 253, 249, 0.68);
  font-size: 1.2rem;
  line-height: 1;
}

.hero__signal > p {
  position: absolute;
  right: -1rem;
  bottom: -0.5rem;
  width: 10rem;
  margin: 0;
  color: var(--color-muted);
  font-family: var(--font-display);
  font-size: 1.55rem;
  line-height: 1.08;
}

.hero__scroll-cue {
  display: flex;
  align-items: center;
  gap: 0.8rem;
  margin-top: auto;
  padding-top: 3.5rem;
  color: var(--color-muted);
  font-size: 0.78rem;
  font-weight: 700;
  letter-spacing: 0.14em;
  text-transform: uppercase;
}

.hero__scroll-cue span {
  color: var(--color-terracotta);
  font-size: 1.4rem;
}

.section--what-is {
  background: var(--color-paper);
}

.section--architecture {
  color: var(--color-paper);
  background: var(--color-ink);
}

.section--team {
  background: var(--color-ivory);
}

.section--contact {
  color: var(--color-paper);
  background: var(--color-sage);
}

.compact-section {
  position: relative;
  z-index: 1;
}

.section-heading {
  max-width: 55rem;
  margin-bottom: clamp(3rem, 6vw, 5.3rem);
}

.section-heading > p:last-child {
  max-width: 39rem;
  margin-bottom: 0;
  color: var(--color-muted);
  font-size: 1.1rem;
}

.section--architecture .section-heading > p:last-child,
.section--contact .section-heading > p:last-child {
  color: rgba(255, 253, 249, 0.72);
}

.section--contact .eyebrow {
  color: var(--color-paper);
  font-size: clamp(1rem, 1.6vw, 1.2rem);
  letter-spacing: 0.2em;
}

.what-is-grid {
  display: grid;
  grid-template-columns: minmax(0, 0.85fr) minmax(0, 1.15fr);
  align-items: start;
  gap: clamp(3rem, 9vw, 9rem);
}

.prose {
  max-width: 42rem;
  color: var(--color-muted);
  font-size: 1.1rem;
}

.prose p:last-child {
  margin-bottom: 0;
}

.principles-list {
  display: grid;
  border-top: 1px solid var(--color-line);
}

.principle {
  display: grid;
  grid-template-columns: 3rem 1fr;
  gap: 0.2rem 1.2rem;
  padding: 1.35rem 0;
  border-bottom: 1px solid var(--color-line);
}

.principle > span,
.plane-card__number,
.flow-step > span {
  color: var(--color-terracotta);
  font-size: 0.76rem;
  font-weight: 700;
  letter-spacing: 0.12em;
  text-transform: uppercase;
}

.principle strong {
  font-size: 1.12rem;
}

.principle p {
  grid-column: 2;
  margin: 0;
  color: var(--color-muted);
  font-size: 0.95rem;
}

.plane-grid {
  display: grid;
  grid-template-columns: repeat(3, minmax(0, 1fr));
  gap: 1px;
  margin-bottom: clamp(3.5rem, 7vw, 6rem);
  background: rgba(255, 253, 249, 0.22);
}

.plane-card {
  min-height: 19rem;
  padding: clamp(1.5rem, 3vw, 2.3rem);
  background: var(--color-ink);
}

.plane-card--featured {
  color: var(--color-paper);
  background: var(--color-terracotta);
}

.plane-card__number {
  display: block;
  margin-bottom: 4.4rem;
  color: rgba(255, 253, 249, 0.67);
}

.plane-card h3 {
  margin-bottom: 1rem;
}

.plane-card p {
  max-width: 23rem;
  margin: 0;
  color: rgba(255, 253, 249, 0.72);
  font-size: 0.97rem;
}

.plane-card--featured p {
  color: rgba(255, 253, 249, 0.84);
}

.architecture-flow {
  display: grid;
  grid-template-columns: minmax(0, 1fr) auto minmax(0, 1fr) auto minmax(0, 1fr) auto minmax(0, 1fr);
  align-items: start;
  gap: 1rem;
  margin-bottom: 3.3rem;
}

.flow-step {
  min-width: 0;
}

.flow-step > span {
  display: block;
  margin-bottom: 1rem;
  color: var(--color-terracotta);
}

.flow-step strong {
  display: block;
  margin-bottom: 0.45rem;
  font-family: var(--font-display);
  font-size: 1.35rem;
  font-weight: 400;
}

.flow-step small {
  display: block;
  color: rgba(255, 253, 249, 0.64);
  font-size: 0.86rem;
  line-height: 1.45;
}

.flow-line {
  padding-top: 1.55rem;
  color: var(--color-terracotta);
  font-size: 1.4rem;
}

.evidence-strip {
  display: grid;
  grid-template-columns: minmax(0, 1.1fr) minmax(0, 0.9fr);
  gap: 2rem;
  padding: 1.45rem 0;
  border-top: 1px solid rgba(255, 253, 249, 0.25);
  border-bottom: 1px solid rgba(255, 253, 249, 0.25);
}

.evidence-strip__label {
  display: block;
  margin-bottom: 0.7rem;
  color: var(--color-terracotta);
  font-size: 0.76rem;
  font-weight: 700;
  letter-spacing: 0.12em;
  text-transform: uppercase;
}

.evidence-strip strong {
  font-family: var(--font-display);
  font-size: clamp(1.15rem, 2vw, 1.55rem);
  font-weight: 400;
}

.evidence-strip p {
  align-self: end;
  margin: 0;
  color: rgba(255, 253, 249, 0.68);
  font-size: 0.95rem;
}

.research-library {
  margin-top: clamp(4rem, 8vw, 6.5rem);
}

.research-library__header {
  display: grid;
  grid-template-columns: minmax(0, 1fr) minmax(16rem, 0.7fr);
  align-items: end;
  gap: 2rem;
  margin-bottom: 1.4rem;
}

.research-library__header h3 {
  max-width: 17ch;
  margin-bottom: 0;
  color: var(--color-paper);
  font-size: clamp(1.8rem, 3vw, 2.7rem);
}

.research-library__header p {
  margin: 0;
  color: rgba(255, 253, 249, 0.68);
  font-size: 0.95rem;
}

.paper-card {
  display: grid;
  grid-template-columns: 10rem minmax(0, 1fr);
  gap: 2rem;
  padding: clamp(1.5rem, 3vw, 2.4rem);
  border: 1px solid rgba(255, 253, 249, 0.28);
  background: var(--color-terracotta);
}

.paper-card__index,
.paper-item__index {
  color: rgba(255, 253, 249, 0.74);
  font-size: 0.76rem;
  font-weight: 700;
  letter-spacing: 0.12em;
  text-transform: uppercase;
}

.paper-card h4 {
  max-width: 34ch;
  margin-bottom: 0.85rem;
  color: var(--color-paper);
  font-family: var(--font-display);
  font-size: clamp(1.5rem, 2.5vw, 2.25rem);
  font-weight: 400;
  line-height: 1.08;
}

.paper-card p {
  max-width: 48rem;
  margin-bottom: 1rem;
  color: rgba(255, 253, 249, 0.82);
}

.paper-card .paper-card__meta,
.paper-item__content .paper-card__meta {
  margin-bottom: 0.9rem;
  color: rgba(255, 253, 249, 0.68);
  font-size: 0.86rem;
}

.paper-card a,
.paper-item__content a {
  color: var(--color-paper);
  font-weight: 700;
  text-decoration: none;
}

.paper-card a:hover,
.paper-card a:focus-visible,
.paper-item__content a:hover,
.paper-item__content a:focus-visible {
  color: #f0c4b8;
}

.paper-list {
  border-top: 1px solid rgba(255, 253, 249, 0.24);
}

.paper-item {
  border-bottom: 1px solid rgba(255, 253, 249, 0.24);
}

.paper-item summary {
  display: grid;
  grid-template-columns: 4rem minmax(0, 1fr) auto;
  align-items: center;
  gap: 1rem;
  padding: 1.3rem 0;
  color: var(--color-paper);
  cursor: pointer;
  list-style: none;
}

.paper-item summary::-webkit-details-marker {
  display: none;
}

.paper-item__index {
  color: var(--color-terracotta);
}

.paper-item__title {
  font-family: var(--font-display);
  font-size: clamp(1.1rem, 1.8vw, 1.45rem);
  line-height: 1.2;
}

.paper-item__toggle {
  color: var(--color-terracotta);
  font-size: 1.5rem;
  line-height: 1;
  transition: transform 180ms ease;
}

.paper-item[open] .paper-item__toggle {
  transform: rotate(45deg);
}

.paper-item__content {
  max-width: 48rem;
  margin-left: 5rem;
  padding: 0 2rem 1.4rem 0;
  color: rgba(255, 253, 249, 0.7);
  font-size: 0.95rem;
}

.paper-item__content p:last-of-type {
  margin-bottom: 0.8rem;
}

.team-group + .team-group {
  margin-top: clamp(3rem, 6vw, 5rem);
}

.team-group__title {
  margin: 0 0 1.25rem;
  color: var(--color-sage);
  font-family: var(--font-body);
  font-size: 0.76rem;
  font-weight: 700;
  letter-spacing: 0.12em;
  line-height: 1.3;
  text-transform: uppercase;
}

.team-profile {
  display: flex;
  min-width: 0;
  flex-direction: column;
  overflow: hidden;
  border: 1px solid var(--color-line);
  background: var(--color-paper);
}

.team-profile--principal {
  display: grid;
  max-width: 54rem;
  grid-template-columns: minmax(16rem, 0.72fr) minmax(0, 1fr);
}

.team-profile__media {
  aspect-ratio: 1 / 1;
  overflow: hidden;
  background: var(--color-soft-gray);
}

.team-profile__media img {
  display: block;
  width: 100%;
  height: 100%;
  object-fit: cover;
  object-position: center 24%;
  filter: grayscale(100%);
}

.team-profile__body {
  display: flex;
  min-height: 10rem;
  flex: 1;
  flex-direction: column;
  padding: 1.5rem;
}

.team-profile--principal .team-profile__body {
  justify-content: center;
  padding: clamp(1.5rem, 4vw, 3rem);
}

.team-profile__role {
  margin: 0 0 0.75rem;
  color: var(--color-terracotta);
  font-size: 0.76rem;
  font-weight: 700;
  letter-spacing: 0.12em;
  line-height: 1.3;
  text-transform: uppercase;
}

.team-profile h3 {
  margin-bottom: 1.5rem;
  font-size: clamp(1.45rem, 2.4vw, 2rem);
  line-height: 1.1;
}

.team-profile__links {
  display: flex;
  flex-wrap: wrap;
  gap: 0.7rem 1rem;
  margin-top: auto;
}

.team-profile__links a {
  color: var(--color-sage);
  font-size: 0.86rem;
  font-weight: 700;
  text-decoration-thickness: 1px;
  text-underline-offset: 0.2em;
}

.team-profile__links a:hover,
.team-profile__links a:focus-visible {
  color: var(--color-terracotta);
}

.team-grid {
  display: grid;
  grid-template-columns: repeat(3, minmax(0, 1fr));
  gap: 1rem;
}

.contact-layout {
  display: grid;
  grid-template-columns: minmax(0, 1fr) minmax(18rem, 0.8fr);
  align-items: end;
  gap: clamp(3rem, 9vw, 8rem);
}

.contact-layout h2 {
  max-width: 10ch;
}

.contact-card {
  padding: 1.6rem;
  border: 1px solid rgba(255, 253, 249, 0.34);
  background: rgba(37, 34, 31, 0.16);
}

.contact-card p {
  margin-bottom: 1.5rem;
  color: rgba(255, 253, 249, 0.8);
  font-size: 1.12rem;
}

.contact-people {
  display: grid;
  gap: 0.8rem;
  margin-bottom: 1.6rem;
}

.contact-person {
  display: flex;
  align-items: baseline;
  justify-content: space-between;
  gap: 1rem;
  padding-top: 0.8rem;
  border-top: 1px solid rgba(255, 253, 249, 0.2);
}

.contact-person__name {
  color: var(--color-paper);
}

.contact-person a {
  color: rgba(255, 253, 249, 0.8);
  font-size: 0.9rem;
  text-decoration-thickness: 1px;
  text-underline-offset: 0.2em;
}

.contact-person a:hover,
.contact-person a:focus-visible {
  color: var(--color-paper);
}

.contact-card .button--quiet {
  border-color: rgba(255, 253, 249, 0.42);
  color: var(--color-paper);
}

.contact-card .button--quiet:hover,
.contact-card .button--quiet:focus-visible {
  border-color: var(--color-paper);
  background: rgba(255, 253, 249, 0.12);
}

.site-footer {
  color: var(--color-muted);
  background: var(--color-ivory);
}

.site-footer__inner {
  display: grid;
  grid-template-columns: auto 1fr auto;
  align-items: center;
  gap: 2rem;
  min-height: 6.5rem;
}

.site-footer p {
  margin: 0;
  font-size: 0.82rem;
  text-align: center;
}

.site-footer .brand {
  color: var(--color-ink);
}

.site-footer__identity {
  gap: 1rem;
}

.institutional-mark--footer {
  font-size: 0.62rem;
}

.anatomy-mark {
  position: absolute;
  z-index: 0;
  pointer-events: none;
  opacity: 0.22;
  animation: anatomy-float 13s ease-in-out infinite;
}

.anatomy-mark path {
  fill: none;
  stroke: var(--color-terracotta);
  stroke-linecap: round;
  stroke-linejoin: round;
  stroke-width: 1.2;
}

.anatomy-mark .anatomy-shade {
  fill: rgba(143, 63, 48, 0.08);
  stroke-width: 0.85;
}

.anatomy-mark .anatomy-hatch {
  stroke: var(--color-sage);
  stroke-width: 0.9;
}

.anatomy-mark--brain {
  top: 8%;
  right: -1rem;
  width: min(31rem, 40vw);
}

.anatomy-mark--thorax {
  top: 12%;
  right: -3rem;
  width: min(29rem, 38vw);
  opacity: 0.25;
}

.section--architecture .anatomy-mark path {
  stroke: #d58673;
}

.section--architecture .anatomy-mark .anatomy-hatch {
  stroke: #83aa95;
}

:focus-visible {
  outline: 3px solid #d58673;
  outline-offset: 4px;
}

html.js-enabled [data-reveal] {
  opacity: 0;
  transform: translateY(1.2rem);
  transition: opacity 620ms ease, transform 620ms ease;
}

html.js-enabled [data-reveal].is-visible {
  opacity: 1;
  transform: none;
}

@keyframes signal-spin-outer {
  from { transform: rotate(0deg); }
  to { transform: rotate(360deg); }
}

@keyframes signal-spin-middle {
  from { transform: rotate(0deg); }
  to { transform: rotate(360deg); }
}

@keyframes signal-spin-inner {
  from { transform: rotate(0deg); }
  to { transform: rotate(360deg); }
}

@keyframes anatomy-float {
  0%, 100% { transform: translate3d(0, 0, 0) rotate(-1deg); }
  50% { transform: translate3d(0, 0.7rem, 0) rotate(1deg); }
}

@media (max-width: 960px) {
  .container {
    width: min(100% - 3.5rem, 760px);
  }

  .hero__grid,
  .what-is-grid,
  .contact-layout {
    gap: 3rem;
  }

  h1 {
    font-size: clamp(3.2rem, 8vw, 5rem);
  }

  .hero__signal {
    width: min(100%, 23rem);
  }

  .site-nav {
    gap: 0.3rem 0.8rem;
  }

  .site-nav a {
    font-size: 0.78rem;
  }

  .anatomy-mark--brain,
  .anatomy-mark--thorax {
    right: -7rem;
    opacity: 0.15;
  }
}

@media (max-width: 760px) {
  .container {
    width: min(100% - 2.5rem, 620px);
  }

  .site-header__inner {
    min-height: 4.9rem;
  }

  .institutional-mark--header {
    display: none;
  }

  .menu-toggle {
    display: grid;
    place-items: center;
  }

  .site-nav {
    position: absolute;
    top: calc(100% + 1px);
    right: 1.25rem;
    left: 1.25rem;
    display: grid;
    gap: 0;
    padding: 0.65rem 1rem;
    border: 1px solid var(--color-line);
    background: var(--color-paper);
    box-shadow: var(--shadow-soft);
    opacity: 0;
    pointer-events: none;
    transform: translateY(-0.5rem);
    transition: opacity 180ms ease, transform 180ms ease;
  }

  .site-nav.is-open {
    opacity: 1;
    pointer-events: auto;
    transform: none;
  }

  .site-nav a {
    padding: 0.8rem 0.25rem;
    border-bottom: 1px solid var(--color-soft-gray);
    font-size: 0.9rem;
  }

  .site-nav a:last-child {
    border-bottom: 0;
  }

  .section {
    padding: 5.5rem 0;
  }

  .section--hero {
    min-height: auto;
    padding-top: 5rem;
    padding-bottom: 2.5rem;
  }

  .hero__grid,
  .what-is-grid,
  .contact-layout {
    grid-template-columns: 1fr;
  }

  .hero__signal {
    width: min(90vw, 22rem);
    margin-top: 1rem;
    margin-bottom: 2rem;
  }

  .hero__signal > p {
    right: -0.6rem;
    bottom: -0.3rem;
    font-size: 1.25rem;
  }

  .hero__scroll-cue {
    padding-top: 2rem;
  }

  h1 {
    max-width: 10ch;
    font-size: clamp(3rem, 14vw, 4.7rem);
  }

  h2 {
    font-size: clamp(2.6rem, 12vw, 4rem);
  }

  .plane-grid,
  .team-grid {
    grid-template-columns: 1fr;
  }

  .plane-card {
    min-height: 15rem;
  }

  .plane-card__number {
    margin-bottom: 2.8rem;
  }

  .architecture-flow {
    grid-template-columns: 1fr;
    gap: 1.6rem;
  }

  .flow-line {
    display: none;
  }

  .evidence-strip {
    grid-template-columns: 1fr;
    gap: 1rem;
  }

  .team-profile--principal {
    grid-template-columns: 1fr;
  }

  .team-profile__body,
  .team-profile--principal .team-profile__body {
    min-height: auto;
    padding: 1.35rem;
  }

  .contact-layout h2 {
    max-width: 12ch;
  }

  .site-footer__inner {
    grid-template-columns: 1fr;
    justify-items: start;
    gap: 1rem;
    padding-top: 2rem;
    padding-bottom: 2rem;
  }

  .site-footer__identity {
    flex-wrap: wrap;
  }

  .site-footer p {
    text-align: left;
  }

  .anatomy-mark--brain,
  .anatomy-mark--thorax {
    top: 4%;
    right: -11rem;
    width: 23rem;
    opacity: 0.1;
  }
}

@media (max-width: 420px) {
  .container {
    width: min(100% - 2rem, 620px);
  }

  .site-header__inner {
    gap: 1rem;
  }

  .brand__name {
    font-size: 1.05rem;
  }

  .brand__mark {
    width: 2.15rem;
    height: 2.15rem;
  }

  .button-row {
    align-items: stretch;
    flex-direction: column;
  }

  .contact-person {
    align-items: flex-start;
    flex-direction: column;
    gap: 0.25rem;
  }

  .button {
    width: 100%;
  }

  .hero__signal {
    width: min(100%, 19rem);
  }

  .hero__signal > p {
    right: -0.4rem;
    width: 8.5rem;
    font-size: 1.1rem;
  }

  .principle {
    grid-template-columns: 2.3rem 1fr;
    gap: 0.2rem 0.8rem;
  }

  .principle p {
    grid-column: 2;
  }

  .research-library__header,
  .paper-card {
    grid-template-columns: 1fr;
  }

  .paper-card__index {
    margin-bottom: -0.5rem;
  }

  .paper-item summary {
    grid-template-columns: 2.4rem minmax(0, 1fr) auto;
    gap: 0.7rem;
  }

  .paper-item__content {
    margin-left: 3.1rem;
    padding-right: 0;
  }
}

@media (prefers-reduced-motion: reduce) {
  *,
  *::before,
  *::after {
    scroll-behavior: auto !important;
    transition-duration: 0.01ms !important;
    animation-duration: 0.01ms !important;
    animation-iteration-count: 1 !important;
  }

  .signal-orbit,
  .anatomy-mark {
    animation: none !important;
  }
}